Hemsleya.
Hemsleya Cogn. ex F. Forbes & Hemsl.
Hemsleya Cogn. ex F. Forbes & Hemsl., J. Linn. Soc. Bot. 23: 490. 1888.
Type: Hemsleya chinensis Cogn. ex F.B. Forbes & Hemsl., A. Henry 2436 (K), China, Hupeh, Patung district.
Perennial climbers or trailers with herbaceous shoots and dioecious sex system. The leaves are simple, petiolate, the blade pedately (3-)5-9(-11)-foliolate. The tendrils are apically bifid or rarely simple. The male flowers are produced in thyrses, female flowers in racemes. The receptacle-tube is rotate with five oblong or lanceolate sepals. The corolla is very variable in form. The five petals are membranaceous, oblong or ovate, white to deep orange-brown. The five stamens consist of monothecous anthers on short, free filaments. The pollen is tricolporate, striate, small (polar axis 33-40 µm, equatorial axis 21-29 µm, (Khunwasi 1998)). The ovary is trilocular at apex and monolocular at the base. The three placentae contain many pendent ovules. The three short styles carry bilobed stigmata. The fruit is a clavate-cylindrical to globose capsule, opening triradiately at the apex. The seeds are compressed with hard testa and usually a woody or rarely membraneous circular wing. The chromosome number is n = 14 (Samuel et al. 1995).
The 28 species are found in China and the eastern Himalayas. Named after the British botanist William Botting Hemsley (1843-1924).
Hemsleya is placed in tribe Gomphogyneae, where it is sister to Gomphogyne (Schaefer et al. 2009; Schaefer & Renner 2011).
Accepted species
Hemsleya amabilis Diels, Notes Roy. Bot. Gard. Edinburgh 5: 206-207. 1912.
Hemsleya carnosiflora C.Y. Wu & Z.L. Chen, Acta Phytotax. Sin. 23: 133-134. 1985.
Hemsleya chengyihana D.Z. Li, Syst. Evol. Hemsleya Cucurbitac.: 91. 1993.
Hemsleya chinensis Cogn. ex F.B. Forbes & Hemsl., J. Linn. Soc., Bot. 23: 490. 1888.
Hemsleya cirromitrata (W.J. de Wilde & Duyfjes) H. Schaef. & S.S. Renner, Taxon 60: 134. 2011.
Hemsleya clavata C.Y. Wu, Acta Phytotax. Sin. 23: 124-125. 1985.
Hemsleya delavayi (Gagnep.) C. Jeffrey ex C.Y. Wu & C.L. Chen, Acta Phytotax. Sin. 23: 134. 1985.
Hemsleya dipterygia Kuang & A.M. Lu, Acta Phytotax. Sin. 20: 88. 1982.
Hemsleya dolichocarpa W.J. Chang, Acta Phytotax. Sin. 21: 190-191. 1983.
Hemsleya dulongjiangensis C.Y. Wu, Acta Phytotax. Sin. 23: 134. 1985.
Hemsleya ellipsoidea L.D. Shen & W.J. Chang, Acta Phytotax. Sin. 21: 185-187. 1983.
Hemsleya emeiensis L.D. Shen & W.J. Chang, Acta Phytotax. Sin. 21: 191. 1983.
Hemsleya endecaphylla C.Y. Wu, Acta Phytotax. Sin. 23: 142. 1985.
Hemsleya gigantha W.J. Chang, Acta Phytotax. Sin. 21: 186-188. 1983.
Hemsleya graciliflora (Harms) Cogn., Pflanzenr. IV, 275 I: 24. 1916.
Hemsleya kunmingensis H.T. Li & D.Z. Li, Ann. Bot. Fenn. 44: 486-487. 2007.
Hemsleya lijiangensis A.M. Lu ex C.Y. Wu & Z.L. Chen, Acta Phytotax. Sin. 23: 129-130. 1985.
Hemsleya longgangensis X.X. Chen & D.R. Liang, Acta Bot. Yunnan. 14: 27-28. 1992.
Hemsleya longivillosa C.Y. Wu & C.L. Chen, Acta Phytotax. Sin. 23: 140. 1985.
Hemsleya macrocarpa (Cogn.) C.Y. Wu ex C. Jeffrey, Kew Bull. 36: 739. 1982.
Hemsleya mitrata C.Y. Wu & Z.L. Chen, Acta Phytotax. Sin. 23: 128-129. 1985.
Hemsleya panacis-scandens C.Y. Wu & Z.L. Chen, Acta Phytotax. Sin. 23: 135-137. 1985.
Hemsleya panlongqi A.M. Lu & W.J. Chang, Acta Phytotax. Sin. 21: 183-185. 1983.
Hemsleya peekelii (W.J. de Wilde & Duyfjes) H. Schaef. & S.S. Renner, Taxon 60: 134. 2011.
Hemsleya pengxianensis W.J. Chang, Acta Phytotax. Sin. 17: 97-98. 1979.
Hemsleya sphaerocarpa Kuang & A.M. Lu, Acta Phytotax. Sin. 20: 87-88. 1982.
Hemsleya turbinata C.Y. Wu, Acta Phytotax. Sin. 23: 140-142. 1985.
Hemsleya zhejiangensis C.Z. Zheng, Acta Phytotax. Sin. 23: 67-68. 1985.
Literature
Khunwasi, C. 1998. Palynology of the Cucurbitaceae. Doctoral Dissertation Naturwiss. Fak., University of Innsbruck.
Li, D.-Z. 1993. Systematics and Evolution of Hemsleya(Cucurbitaceae). Yunnan: Yunnan Science and Technology Press. 126 pp., 32 plates. [in Chinese with English summary]
Samuel, R., Balasubramaniam, S., and W. Morawetz. 1995. The karyology of some cultivated Cucurbitaceae of Sri Lanka. Ceylon J. Sci., Biol. Sci. 24: 17-22.
Schaefer, H. and S.S. Renner. 2011. Phylogenetic relationships in the order Cucurbitales and a new classification of the gourd family (Cucurbitaceae). Taxon 60: 122-138.
Schaefer, H., Heibl, C., and S.S. Renner. 2009. Gourds afloat: a dated phylogeny reveals an Asian origin of the gourd family (Cucurbitaceae) and numerous oversea dispersal events. Proceedings of the Royal Society B 276: 843-851.